A feature of these tyres is, they have spokes with a special curve arranged in an interlacing fashion.
http://www.national.co.uk/press-releases/123/bridgestone-announces-development-of-non-pneumatic--airless--concept-tyre/
http://www.national.co.uk/branch-513-Blackburn.aspx
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Blackburn
BB2 2DH
This enables them to flex effectively without twisting, and behave just like ordinary tyres, without producing a horizontal force.